Portal:Amphibians
The Amphibian Portal
Amphibians are ectothermic, anamniotic, four-limbed vertebrate animals that constitute the class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all tetrapods, excluding the amniotes (tetrapods with an amniotic membrane, such as modern reptiles, birds, and mammals). All extant (living) amphibians belong to the monophyletic subclass Lissamphibia, with three living orders: Anura (frogs), Urodela (salamanders), and Gymnophiona (caecilians). Evolved to be mostly semiaquatic, amphibians have adapted to inhabit a wide variety of habitats, with most species living in freshwater, wetland or terrestrial ecosystems (such as riparian woodland, fossorial and even arboreal habitats). Their life cycle typically starts out as aquatic larvae with gills known as tadpoles, but some species have developed behavioural adaptations to bypass this.
The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards, but unlike reptiles and other amniotes, require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in recent decades there has been a dramatic decline in amphibian populations for many species around the globe.
The earliest amphibians evolved in the Devonian period from sarcopterygian fish with lungs and bony-limbed fins, features that were helpful in adapting to dry land. They diversified and became ecologically dominant during the Carboniferous and Permian periods, but were later displaced in terrestrial environments by early reptiles and basal synapsids (mammal predecessors). The origin of modern amphibians belonging to Lissamphibia, which first appeared during the Early Triassic, around 250 million years ago, has long been contentious. The most popular hypothesis is that they likely originated from temnospondyls, the most diverse group of prehistoric amphibians, during the Permian period. Another hypothesis is that they emerged from lepospondys. (Full article...)
Selected frog article
Selected salamander article
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Sexual selection in amphibians involves sexual selection processes in amphibians, including frogs, salamanders and newts. Prolonged breeders, the majority of frog species, have breeding seasons at regular intervals where male-male competition occurs with males arriving at the waters edge first in large number and producing a wide range of vocalizations, with variations in depth of calls the speed of calls and other complex behaviours to attract mates. The fittest males will have the deepest croaks and the best territories, with females making their mate choices at least partly based on the males depth of croaking. This has led to sexual dimorphism, with females being larger than males in 90% of species, males in 10% and males fighting for groups of females.
There is a direct competition between males to win the attention of the females in salamanders and newts, with elaborate courtship displays to keep the females attention long enough to get her interested in choosing him to mate with. Some species store sperm through long breeding seasons, as the extra time may allow for interactions with rival sperm. (Full article...)List of selected salamander articles
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Did you know? –
- ... that the giant spiny frog is the largest frog in Hong Kong?
- ... that where their ranges overlap, the Italian cave salamander sometimes hybridises with Ambrosi's cave salamander?
- ... that Pelobates cultripes, the "Western Spadefoot Toad", produce spawn up to one metre long which can consist of 7000 eggs?
- ... that the shovelnose salamander and the blackbelly salamander both live in the Appalachian Mountains, but the former feeds exclusively in water while the latter mainly forages on land?
- ... that the large prehistoric amphibian Cyclotosaurus had a skull up to 70 cm long?
Selected amphibian type
Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All ten extant salamander families are grouped together under the order Urodela from the group Caudata. Urodela is a scientific Latin term based on the Ancient Greek οὐρά δήλη: ourà dēlē "conspicuous tail". Caudata is the Latin for "tailed ones", from cauda : "tail".
Salamander diversity is highest in eastern North America, especially in the Appalachian Mountains; most species are found in the Holarctic realm, with some species present in the Neotropical realm. Salamanders never have more than four toes on their front legs and five on their rear legs, but some species have fewer digits and others lack hind limbs. Their permeable skin usually makes them reliant on habitats in or near water or other cool, damp places. Some salamander species are fully aquatic throughout their lives, some take to the water intermittently, and others are entirely terrestrial as adults.
This group of amphibians is capable of regenerating lost limbs as well as other damaged parts of their bodies. Researchers hope to reverse engineer the regenerative processes for potential human medical applications, such as brain and spinal cord injury treatment or preventing harmful scarring during heart surgery recovery. The remarkable ability of salamanders to regenerate is not just limited to limbs but extends to vital organs such as the heart, jaw, and parts of the spinal cord, showing their uniqueness compared to different types of vertebrates. This ability is most remarkable for occurring without any type of scarring. This has made salamanders an invaluable model organism in scientific research aimed at understanding and achieving regenerative processes for medical advancements in human and animal biology. (Full article...)Selected images
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Photo credit: LiquidGhoulThe New England Tree Frog (Litoria subglandulosa) is a species of frog native to the streams of the New England Tablelands, New South Wales, Australia. -
Photograph: Benny TrappThe Iberian midwife toad (Alytes cisternasii) is a species of frog in the family Alytidae found in Portugal and western Spain. Its natural habitats are temperate forests and shrubland, Mediterranean-type shrubby vegetation, intermittent rivers and freshwater marshes, pastureland, and aquaculture ponds. The species is affected by habitat loss, and the IUCN lists it as being "Near Threatened". -
Photograph: Benny TrappThe Epirus water frog (Pelophylax epeiroticus) is a species of frog in the family Ranidae. It is found in western Greece, including Corfu, and the southern areas of Albania. The species is collected from the wild for human consumption. -
The Blue Mountains Tree Frog (Litoria citropa) is a moderate-sized species of tree frog, up to about 60 mm (2.4 in) in length. It is native to coastal and highland areas of eastern Australia, especially in the Blue Mountains, hence its name. -
The edible frog (Pelophylax kl. esculentus) is a fertile hybrid of the pool and marsh frogs which is commonly found in Europe. The species is commonly used in food, including the French delicacy frog legs. -
The emerald-eyed tree frog (Rana platanera) is a species of frog in the family Hylidae found in South America. -
Photo: Marie-Lan NguyenThe marsh frog (Pelophylax ridibundus) is a species of true frog and the largest frog native to Europe; females of this sexually dimorphic species may be up to 17 centimetres (6.7 in) long. The marsh frog feeds mainly on insects, but it also eats smaller amphibians, fish, and rodents. -
Photograph: Brian GratwickeRabbs' fringe-limbed treefrog (Ecnomiohyla rabborum) is a large species of frog originally found in the forest canopies of central Panama. Only discovered in 2005, the species is thought to be extinct in the wild; only one specimen, a male at the Atlanta Botanical Garden, survives. -
Photograph: Benny TrappThe Spanish painted frog (Discoglossus jeanneae) is a species of frog in the family Alytidae. Endemic to Spain, it mostly lives in open areas, pine groves and shrublands. It feeds mostly on insects and worms. -
Photo credit: Patrick CoinA terrestrial subadult Eastern newt or red eft (Notophthalmus viridescens). Salamanders of the family Salamandridae with aquatic adult stages are called newts. Some newts, including the Eastern newt, have a juvenile terrestrial stage called the eft. The red eft has bright aposematic coloration to warn predators of its highly toxic skin. -
The white-lipped tree frog (Litoria infrafrenata) is the world's largest tree frog. This species is native to the rainforests of Northern Queensland, New Guinea, the Bismarck Islands and the Admiralty Islands. It can reach a length of over 13 cm (5 in), with females growing larger than males. The lower lip has a distinctive white stripe, giving this species its common name. -
Atelopus certus is a species of toad endemic to the Darién Province of eastern Panama. Much of its range falls within the Darién National Park, a protected area and World Heritage Site. It is classified as endangered, primarily due to the advancing wave of chytridiomycosis moving through Central America. -
Photograph credit: RushenbWallace's flying frog (Rhacophorus nigropalmatus) is a moss frog found in tropical southeastern Asia. It is named after the British naturalist Alfred R. Wallace, who collected the first known specimen of the species. It lives almost exclusively in trees, and when threatened, or in search of prey, will leap from a branch and splay its four webbed feet; the membranes between its toes and the loose skin flaps on its sides catch the air as it falls, helping it to glide. This individual was photographed in Khao Sok National Park, Thailand. -
Photograph credit: RushenbRaorchestes parvulus is a small bush frog native to tropical southeastern Asia. Although the reproductive strategy of this species has not been studied, members of the family typically make a foam nest while mating in vegetation overhanging a stream. They create the foam by beating their legs, and the eggs are deposited into the nest and covered with seminal fluid before the foam hardens into a protective casing. When the eggs hatch, the young push their way out of the nest and fall into the water below.
This picture shows a male R. parvulus frog, photographed in Phu Kradueng National Park, Thailand, calling by inflating its vocal sac. -
Photo: LokionlyThe Common Indian Toad (Duttaphrynus melanostictus) is native to South Asia. It grows to about 20 cm (8 in) long and breeds during monsoon season. These toads are often seen at night under street lamps especially during times when winged termites swarm. They have been noted to feed on a wide range of invertebrates, including scorpions.
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Selected toad article
The neural basis of prey detection, recognition, and orientation was studied in depth by Jörg-Peter Ewert in a series of experiments that made the toad visual system a model system in neuroethology (neural basis of natural behavior). He began by observing the natural prey catching behavior of the common European toad (Bufo bufo).
Ewert's work with toads yielded several important discoveries (Ewert 1974, 2004). In general, his research revealed the specific neural circuits for recognition of complex visual stimuli. Specifically, he identified two main regions of the brain, the tectum and the thalamic-pretectal region, that were responsible for discriminating prey from non-prey and revealed the neural pathways that connected them. Furthermore, he found that the neural mechanisms are plastic and adaptable to varying environments and conditions (Carew 2000; Zupanc 2004). (Full article...)Selected caecilian article
Typhlonectidae, also known as aquatic caecilians or rubber eels, are a family of caecilians found east of the Andes in South America.
They are viviparous animals, giving birth to young that possess external gills. Of the five extant genera in the family, Atretochoana, Potamotyphlus and Typhlonectes are entirely aquatic, while Chthonerpeton and Nectocaecilia are semi-aquatic. Atretochoana reaches 100 cm (39 in) in length, but other species in the family range from 20 to 60 cm (7.9–23.6 in). A sixth typhlonectid genus, Ymboirana, was recently described and is known exclusively from fossil material. (Full article...)Need help?
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